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Chalcosoma caucasus (page 7)

In the scenes below are shown two males engaged in battle.  These fights are carried out over breeding rights or to defend feeding sites.  Such encounters do not usually end in serious injury.  However, due their enormous strength, they are quite capable of piercing each other's armor with their pointed horns.  Additionally, this species has a razor sharp edge on the thoracic shield or "pronotum".  This edge can be pressed firmly against the area where it meets the elytra, and used to snap off an opponent's leg, or ward off an unwary predator.  As can be seen in some of the images, a male will occasionally succeed in prying its opponent from its foothold, and the rival will be lifted completely off the ground.

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Chalcosoma caucasus males - Image  C. Campbell
Chalcosoma caucasus males - Image  C. Campbell

 
Chalcosoma caucasus males - Image  C. Campbell
Chalcosoma caucasus males - Image  C. Campbell

 
Chalcosoma caucasus males - Image  C. Campbell
Chalcosoma caucasus males - Image  C. Campbell

 
Chalcosoma caucasus males - Image  C. Campbell
Chalcosoma caucasus males - Image  C. Campbell

 
Chalcosoma caucasus males - Image  C. Campbell

 
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