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BIOLOGY:
- ANATOMY -
INTERNAL ANATOMY (page 6)
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The Internal Anatomy of the Thylacine - A Historical Perspective

   Professor James Peter Hill (1873-1954), was an eminent embryologist who specialised in the study of monotremes and marsupials.  Born and educated in Scotland, he became a demonstrator of biology at the University of Sydney in 1892, and Lecturer on Embryology in 1904.  Hill went on to become Professor of Embryology at University College in London from 1921 to 1938.  A biographical summary of Hill's career published in the Journal of Anatomy (Anon. 1948) states:

    "It may be asserted without danger of exaggeration that J. P. Hill has not only laid the foundations of our knowledge of the development of Monotremes and Marsupials, but that he has, by his own researches and those which he has stimulated others to undertake, erected thereon a lasting monument of exact knowledge which constitutes a contribution to science of permanent and inestimable value".

    Hill dissected the reproductive tract of a female thylacine sent to him by Alexander Morton, the curator of the Royal Society of Tasmania's Museum, on the 5th August 1902.  Although no publication resulted from this endeavour, Hill's working notebook, together with his dissection, survives in the collection of the Museum für Naturkunde (Humboldt University) in Berlin.  Hill's other legacy, also within the museum's collection, is a series of stained slides of the reproductive anatomy of the female thylacine, which number some 200 in total.  The slides detail the right and left ovaries, utero-tubal junction, infundibulum of the fallopian tube, os uterus and cervix (Source: ITSD 5th Revision 2013).

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preserved reproductive tract of the female thylacine
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Reproductive tract of female thylacine (Specimen MA 840).  Museum für Naturkunde (Humboldt University), Berlin.  Source: ITSD 5th Revision 2013.
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    Sir Colin MacKenzie (1877-1938) was an eminent, Melbourne-based orthopaedic surgeon who devoted much of his life to the study of Australian fauna.  In 1919, he formed and financed the Australian Institute of Anatomical Research, and began building a collection of preserved specimens of Australian wildlife.  This work intensified in the 1920s when the Victorian government granted him permission to establish a field research station at Healesville that enabled him to breed and collect native animals for use as anatomical specimens.
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entrance gates to Sir Colin McKenzie Sanctuary
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Entrance gates to Sir Colin MacKenzie Sanctuary at Healesville (Victoria).  Courtesy: State Library of Victoria.
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    In the course of his work, MacKenzie accumulated the largest single collection of thylacine organ specimens in existence (Source: ITSD 5th Revision 2013).  In the Age newspaper of the 13th August 1925 (p. 13), MacKenzie wrote:

    "Unfortunately these animals are fast disappearing, and, in less than twenty years it is computed, will, in the absence of rigid protective measures, be all extinct".

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Australian Institute of Anatomy - 1938
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A photo taken in 1938 of the Australian Institute of Anatomy in Canberra, where the vast collection of anatomical specimens (including those of the thylacine) accumulated by Sir Colin MacKenzie were once housed.  Amongst the institute's functions were serving as a natural history museum and conducting nutritional research.
    MacKenzie generously donated his entire collection of marsupial specimens, including those of the thylacine, to the Australian nation and in 1924 the government responded by creating the National Museum of Australian Zoology to house them, appointing him as its first director.  In 1931, the museum became known as the Australian Institute of Anatomy to coincide with the opening of its Canberra home.  The Australian Institute of Anatomy closed in December 1985 and the MacKenzie collection transferred to its current home at the National Museum of Australia in Canberra.

    MacKenzie's focus on the anatomical attributes of his specimens reduced their value from a zoological perspective as he failed to keep detailed records for each specimen, only labelling them with a brief description of the organ preserved and the common name of the animal (Source: ITSD 5th Revision 2013).  Although MacKenzie did not contribute directly to our knowledge of the internal anatomy of the thylacine, his legacy was the foresight to collect and preserve all of its organ systems for future generations of scientists to study.

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References
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