Because some taxonomists of the nineteenth century failed to recognize
the variations which occur within the species
T. cynocephalus, the
classification of fossil thylacinids has been complicated. For example,
when DeVis (1894) received the left side of a skull discovered at Ellagowan,
on the Darling Downs, he made a comparison between it and only a single
skull of the modern species. Finding the fossilized skull to be somewhat
larger, he declared it to be that of a separate species,
Thylacinus
rostralis. Thylacinus breviceps was assumed to be a small
species, but was actually based upon an immature specimen (Moeller 1968).
However, DeVis (1894) realized that T. major from the Wellington
Caves was identical to the widespread "cave tiger",
T spelaeus,
which Ride (1964) demonstrated to be the same as the modern species.
Ride (1964) studied both fossil and recent examples of thylacine material
from throughout Australia. He was able to show that T. spelaeus
falls within the range of variation of T. cynocephalus by taking
eight measurements on each specimen.
.
| Plate
5 from Richard Owen's (1877) "Researches on the fossil remains of the
extinct mammals of Australia..." which depicts fossils from the caves
of New South Wales. Included are some illustrations (second row of
figures from the bottom) of Thylacinus spelaeus (=cynocephalus). |
|
However, he did not make a conclusive decision on T. rostralis,
commenting that the specimens found in the southwestern region of the continent
appeared to be smaller than average. To avoid the complications which
can occur when measuring juvenile skulls, Lowry (1972) took measurements
of the specimens' teeth, which do not grow continuously throughout life.
The fossils from the southwest were 10% smaller than the average, and ones
from Eucla 13% smaller, but were still within the acceptable size range
for a single species. Therefore, based upon current evidence, all
thylacine material of Quaternary age (Pleistocene and Holocene epochs)
should be considered as belonging to the modern species. However,
Thylacinus
potens, the Late Miocene species described by Woodburne
(1967), possesses anatomical differences which distinguish it from the
modern T. cynocephalus, most notably, in the structure of its palate
and molar teeth. From the Alcoota Local Fauna of the Northern Territory,
T.
potens was also more massive in body build than the modern thylacine. |
In Tasmania, T. cynocephalus
fossils have been found at Scotchtown (Gill 1953). In Victoria, fossil
material has been found at Keilor (Marshall 1974), in a cave near Gismore
(Gill 1953), at Buchan (Hope 1974), Mt. Hamilton, 175 km west of Melbourne
(Wakefield 1963), and at Nelson (Colliver 1938). In South Australia,
specimens of Pleistocene age were discovered within Victoria
Cave, near Naracoorte (Smith 1972). At nearby Henschke's
Quarry Cave, more Pleistocene specimens were
found which are around 32,000 years old (Pledge 1974), and occur in association
with the fossils of species that are characteristic of the dry sclerophyll
forests and dense heath habitats which still exist in the region today.
Additional South Australian localities for thylacine material include the
Buckalowie Caves near Carrieton (Jones 1923), and an isolated, malformed
tooth from Fromm's Landing, dated at 3,800 BP (Mulvaney et. al.
1964, and Archer 1971).
In New South Wales,
thylacine fossils have been found in the Wellington
Caves (Owen 1877), the Wombeyan Caves (Broom 1896), and Lake Menindee
(Tedford 1967). The Lake Menindee specimens were recovered from a
stratum between two layers of charcoal, dated at 26,300 and 18,800 years
BP respectively. In Queensland, sites include Cement Mills, Gore
(Bartholomai 1977), Ellangowan, Pilton, Gowrie Creek and Chinchilla (DeVis
1894).
.
| A
map of significant localities of Quaternary (Pleistocene and Holocene epochs)
mammal remains in Australasia, a number of which are mentioned in the text
(after Archer et al. 1984) |
| 1.
Tari |
| 2. Nombe
& Kiowa |
| 3. Glen
Garland |
| 4. Chillagoe
Caves & Tea Tree Cave |
| 5. Herveys
Range |
| 6. Marmor |
| 7. East
Darling Downs |
| 8. Gore |
| 9. Russenden
Cave & The Joint |
| 10. Cuddie Springs |
| 11. Bingara |
| 12. Murrurundi |
| 13. Scone |
| 14. Mendooran |
| 15. Wellington Caves |
| 16. Douglas Cave |
| 17. Wombeyan Caves |
| 18. Lake George |
| 19. Mount Fairy |
| 20. Cloggs Cave &
Pyramids Cave |
| 21. Brighton |
| 22. Queenscliff |
| 23. Maribyrnong |
| 24. Lancefield |
|
| 25. Kow Swamp |
| 26. Loddon River |
| 27. Waurn Ponds |
| 28. Colac |
| 29. Torquay |
| 30. Lake Colongulac |
| 31. Ulverstone |
| 32. Scotchdown Cave |
| 33. Mowbray Swamp |
| 34. Pleisto Scene
Cave |
| 35. Porteus Spring
Mound |
| 36. Port Fairy |
| 37. Portland |
| 38. Spring Creek |
| 39. Grange Burn &
Muddy Creek |
| 40. McEachern's Cave |
| 41. Mount Gambier
Caves |
| 42. Green Waterhole |
| 43. Victoria Cave |
| 44. Fromm's landing |
| 45. Lake Victoria
& Frenchman's Creek |
| 46. Lake Mungo |
| 47. Tandou |
| 48. Menindee |
|
| 49. Bileroo Creek |
| 50. Yanda |
| 51. Lake Callabonna |
| 52. Malkuni |
| 53. Kanunka |
| 54. Cooper Creek |
| 55. Warburton River |
| 56. Hookina Creek |
| 57. Buckalowie Caves |
| 58. Lake Fowler |
| 59. Seton Rock Shelter |
| 60. Rocky River |
| 61. Koonalda Cave |
| 62. Thylacine Hole |
| 63. Horseshoe Cave |
| 64. Madura Cave |
| 65. Murra-el-elelevyn
Cave |
| 66. Balladonia |
| 67. Devil's Lair |
| 68. Labyrinth Cave |
| 69. Mammoth Cave |
| 70. Hastings Cave |
| 71. Murchison River |
| 72. Cape Range Caves |
|
|