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THE NATURAL HISTORY OF THYLACINUS CYNOCEPHALUS:
- PREHISTORIC RANGE OF THE THYLACINE -
(page 2)
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    Because some taxonomists of the nineteenth century failed to recognize the variations which occur within the species T. cynocephalus, the classification of fossil thylacinids has been complicated.  For example, when DeVis (1894) received the left side of a skull discovered at Ellagowan, on the Darling Downs, he made a comparison between it and only a single skull of the modern species.  Finding the fossilized skull to be somewhat larger, he declared it to be that of a separate species, Thylacinus rostralis.  Thylacinus breviceps was assumed to be a small species, but was actually based upon an immature specimen (Moeller 1968).  However, DeVis (1894) realized that T. major from the Wellington Caves was identical to the widespread "cave tiger", T spelaeus, which Ride (1964) demonstrated to be the same as the modern species.  Ride (1964) studied both fossil and recent examples of thylacine material from throughout Australia.  He was able to show that T. spelaeus falls within the range of variation of T. cynocephalus by taking eight measurements on each specimen.
 
Pleistocene marsupial fossils
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Plate 5 from Richard Owen's (1877) "Researches on the fossil remains of the extinct mammals of Australia..." which depicts fossils from the caves of New South Wales.  Included are some illustrations (second row of figures from the bottom) of Thylacinus spelaeus (=cynocephalus).
    However, he did not make a conclusive decision on T. rostralis, commenting that the specimens found in the southwestern region of the continent appeared to be smaller than average.  To avoid the complications which can occur when measuring juvenile skulls, Lowry (1972) took measurements of the specimens' teeth, which do not grow continuously throughout life.  The fossils from the southwest were 10% smaller than the average, and ones from Eucla 13% smaller, but were still within the acceptable size range for a single species.  Therefore, based upon current evidence, all thylacine material of Quaternary age (Pleistocene and Holocene epochs) should be considered as belonging to the modern species.  However, Thylacinus potens, the Late Miocene species described by Woodburne (1967), possesses anatomical differences which distinguish it from the modern T. cynocephalus, most notably, in the structure of its palate and molar teeth.  From the Alcoota Local Fauna of the Northern Territory, T. potens was also more massive in body build than the modern thylacine.

    In Tasmania, T. cynocephalus fossils have been found at Scotchtown (Gill 1953).  In Victoria, fossil material has been found at Keilor (Marshall 1974), in a cave near Gismore (Gill 1953), at Buchan (Hope 1974), Mt. Hamilton, 175 km west of Melbourne (Wakefield 1963), and at Nelson (Colliver 1938).  In South Australia, specimens of Pleistocene age were discovered within Victoria Cave, near Naracoorte (Smith 1972).  At nearby Henschke's Quarry Cave, more Pleistocene specimens were found which are around 32,000 years old (Pledge 1974), and occur in association with the fossils of species that are characteristic of the dry sclerophyll forests and dense heath habitats which still exist in the region today.  Additional South Australian localities for thylacine material include the Buckalowie Caves near Carrieton (Jones 1923), and an isolated, malformed tooth from Fromm's Landing, dated at 3,800 BP (Mulvaney et. al. 1964, and Archer 1971).

    In New South Wales, thylacine fossils have been found in the Wellington Caves (Owen 1877), the Wombeyan Caves (Broom 1896), and Lake Menindee (Tedford 1967).  The Lake Menindee specimens were recovered from a stratum between two layers of charcoal, dated at 26,300 and 18,800 years BP respectively.  In Queensland, sites include Cement Mills, Gore (Bartholomai 1977), Ellangowan, Pilton, Gowrie Creek and Chinchilla (DeVis 1894).
 

Localities of Quaternary mammal remains in Australasia - image © C. Campbell
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A map of significant localities of Quaternary (Pleistocene and Holocene epochs) mammal remains in Australasia, a number of which are mentioned in the text (after Archer et al. 1984)
 
 
 1.   Tari
 2.   Nombe & Kiowa
 3.   Glen Garland
 4.   Chillagoe Caves & Tea Tree Cave
 5.   Herveys Range
 6.   Marmor
 7.   East Darling Downs
 8.   Gore
 9.   Russenden Cave & The Joint
10.  Cuddie Springs
11.  Bingara
12.  Murrurundi
13.  Scone
14.  Mendooran
15.  Wellington Caves
16.  Douglas Cave
17.  Wombeyan Caves
18.  Lake George
19.  Mount Fairy
20.  Cloggs Cave & Pyramids Cave
21.  Brighton
22.  Queenscliff
23.  Maribyrnong
24.  Lancefield
 
25.  Kow Swamp
26.  Loddon River
27.  Waurn Ponds
28.  Colac
29.  Torquay
30.  Lake Colongulac
31.  Ulverstone
32.  Scotchdown Cave
33.  Mowbray Swamp
34.  Pleisto Scene Cave
35.  Porteus Spring Mound
36.  Port Fairy
37.  Portland
38.  Spring Creek
39.  Grange Burn & Muddy Creek
40.  McEachern's Cave
41.  Mount Gambier Caves
42.  Green Waterhole
43.  Victoria Cave
44.  Fromm's landing
45.  Lake Victoria & Frenchman's Creek
46.  Lake Mungo
47. Tandou
48. Menindee
 
49.  Bileroo Creek
50.  Yanda
51.  Lake Callabonna
52.  Malkuni
53.  Kanunka
54.  Cooper Creek
55.  Warburton River
56.  Hookina Creek
57.  Buckalowie Caves
58.  Lake Fowler
59.  Seton Rock Shelter
60.  Rocky River
61.  Koonalda Cave
62.  Thylacine Hole
63.  Horseshoe Cave
64.  Madura Cave
65.  Murra-el-elelevyn Cave
66.  Balladonia
67.  Devil's Lair
68.  Labyrinth Cave
69.  Mammoth Cave
70.  Hastings Cave
71.  Murchison River
72.  Cape Range Caves
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Information on this page is referenced primarily from:  SMITH, M., 1982. Review of the Thylacine (Marsupialia, Thylacinidae). In "Carnivorous Marsupials - Vol. 1" (Ed. M. Archer). Roy. Zool. Soc. N.S.W.: Sydney. pp. 237-53.
Section references
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